Life cycle on barberry plant. Here spermatization, pyniospores and aeciospores have been described. #puccinia #secondaryhost #barberry #basidiospore #pycnidiospore #aeciospore #aeciodiospore #aecidialcup #pycnialcup #spermatization #ostiole #receptivehypha #flexuoushyphae #dikaryotization